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The flightless dung beetle (Circellium bacchus) is a species of dung beetle endemic to a few areas of South Africa, including the Addo Elephant National Park and the Kariega Game Reserve where this image was taken. The species was originally widespread in Southern Africa, but it only survives in the few areas mentioned above; as such, it qualifies as a IUCN vulnerable species. Its vulnerability is exacerbated by a number of other factors, including the fact that its habitat is under threat by agriculture and human activity, that is has low breeding capacity as well as low dispersability (as a consequence of being flightless), and that its survival is strictly dependent on a number of vertebrates (particulary elephant and buffalo) that are also experiencing a decrease in population.
The flightless dung beetles mostly feed on elephant or buffalo faeces, but they have been recorded to also feed on dung from other species such as rabbits, baboons, antelopes, and ostrichs.